Dubiously lichenized ascomycete from Denmark

This miniscule species was found overgrowing mosses and detritus at a small abandoned piece of land that was once a part a car mechanic workshop, next to a heavily trafficed road in Aarhus, Denmark. At first glance it appears clearly lichenized with a somewhat gelatinous thallus, but the photobiont is very difficult to determine in sections. There are some quite big globular structures with a multi-layered wall in the matrix that could be zygospores or maybe akinetes and some of them may be penetrated by hyphae. Nostoc is also present in the matrix as are a few chlorococcoid algae and a filamentous alga bearing some resemblance to Mougeotia which in turn could be the source of the “zygospores”.

The largest apothecia measure around 0.2 mm across, appear pencil-like when immature (like Jamesiella-hyphophores) becoming short-stipitate and +/- urceolate with age. The color ranges from whitish to pale pink to orangeish in the oldest (senescent) apothecia. Excipulum looks poorly developed, consisting of what appears to be a single layer of +/- fused parallel hyphae. Asci very easily detached from hypothecium, moderately thickwalled with a clear apical expansion that by the looks of it lacks a tholus. Apical expansion with a K/I+ blue cushion penetrated by a tube-like structure – I think one of the ascus walls is weakly K/I+ blue as well. Paraphyses long and slender not or perhaps weakly anastomosing and weakly expanded apically. Spores 8/ascus, containing many small droplets, (0-)1-3 septate, ~24-26 µm x 9-11 µm, the most mature-looking ones slightly constricted at septae. I inquired about this collection in the facebook-group “Lichens Connecting People” where no suggestion as to the ID of the species was given, but the septation of the spores proved to be a point of contention as some of them appear (sub-)muriform on photographs, but it is my impression when observing them under the microscope that it is incompletely coalesced lipid droplets making them appear this way.

I considered Coppinsia minutissima, but spore-size and septation would rule out that species, however the morphology and ascus-struture looks so distinct that I am hoping that one of you might have an idea as to the correct ID. Any and all suggestions and comments are most welcome.

-Jonas

Edit: I forgot the external link restriction and removed link to flickr, but my uploaded photos don’t seem to show up here. Working on remedying this

Photos:

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